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The Werner syndrome protein promotes CAG/CTG repeat stability by resolving large (CAG)(n)/(CTG)(n) hairpins.
[werner syndrome]
Expansion
of
CAG
/
CTG
repeats
causes
certain
neurological
and
neurodegenerative
disorders
,
and
the
formation
and
subsequent
persistence
of
stable
DNA
hairpins
within
these
repeats
are
believed
to
contribute
to
CAG
/
CTG
repeat
instability
.
Human
cells
possess
a
DNA
hairpin
repair
(
HPR
)
pathway
,
which
removes
various
(
CAG
)
(
n
)
and
(
CTG
)
(
n
)
hairpins
in
a
nick
-directed
and
strand-
specific
manner
.
Interestingly
,
this
HPR
system
processes
a
(
CTG
)
(
n
)
hairpin
on
the
template
DNA
strand
much
less
efficiently
than
a
(
CAG
)
(
n
)
hairpin
on
the
same
strand
(
Hou
,
C
.
,
Chan
,
N
.
L
.
,
Gu
,
L
.
,
and
Li
,
G
.
M
.
(
2009
)
Incision-dependent
and
error-free
repair
of
(
CAG
)
(
n
)
/
(
CTG
)
(
n
)
hairpins
in
human
cell
extracts
.
Nat
.
Struct
.
Mol
.
Biol
.
16
,
869
-
875
)
,
suggesting
the
involvement
of
an
additional
component
for
(
CTG
)
(
n
)
HPR
.
To
identify
this
activity
,
a
functional
in
vitro
HPR
assay
was
used
to
screen
partially
purified
HeLa
nuclear
fractions
for
their
ability
to
stimulate
(
CTG
)
(
n
)
HPR
.
We
demonstrate
here
that
the
stimulating
activity
is
the
Werner
syndrome
protein
(
WRN
)
.
Although
WRN
contains
both
a
3
'
→
5
'
helicase
activity
and
a
3
'
→
5
'
exonuclease
activity
,
the
stimulating
activity
was
found
to
be
the
helicase
activity
,
as
a
WRN
helicase
mutant
failed
to
enhance
(
CTG
)
(
n
)
HPR
.
Consistently
,
WRN
efficiently
unwound
large
(
CTG
)
(
n
)
hairpins
and
promoted
DNA
polymerase
δ-catalyzed
DNA
synthesis
using
a
(
CTG
)
(
n
)
hairpin
as
a
template
.
We
,
therefore
,
conclude
that
WRN
stimulates
(
CTG
)
(
n
)
HPR
on
the
template
DNA
strand
by
resolving
the
hairpin
so
that
it
can
be
efficiently
used
as
a
template
for
repair
or
replicative
synthesis
.
Diseases
Validation
Diseases presenting
"specific manner"
symptom
aromatase deficiency
esophageal adenocarcinoma
esophageal carcinoma
familial hypocalciuric hypercalcemia
gm1 gangliosidosis
holt-oram syndrome
werner syndrome
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