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Acetylation of Werner syndrome protein (WRN): relationships with DNA damage, DNA replication and DNA metabolic activities.
[werner syndrome]
Loss
of
Werner
syndrome
protein
function
causes
Werner
syndrome
,
characterized
by
increased
genomic
instability
,
elevated
cancer
susceptibility
and
premature
aging
.
Although
WRN
is
subject
to
acetylation
,
phosphorylation
and
sumoylation
,
the
impact
of
these
modifications
on
WRN
's
DNA
metabolic
function
remains
unclear
.
Here
,
we
examined
in
further
depth
the
relationship
between
WRN
acetylation
and
its
role
in
DNA
metabolism
,
particularly
in
response
to
induced
DNA
damage
.
Our
results
demonstrate
that
endogenous
WRN
is
acetylated
somewhat
under
unperturbed
conditions
.
However
,
levels
of
acetylated
WRN
significantly
increase
after
treatment
with
certain
DNA
damaging
agents
or
the
replication
inhibitor
HU
.
Use
of
DNA
repair-
deficient
cells
or
repair
pathway
inhibitors
further
increase
levels
of
acetylated
WRN
,
indicating
that
induced
DNA
lesions
and
their
persistence
are
at
least
partly
responsible
for
increased
acetylation
.
Notably
,
acetylation
of
WRN
correlates
with
inhibition
of
DNA
synthesis
,
suggesting
that
replication
blockage
might
underlie
this
effect
.
Moreover
,
WRN
acetylation
modulates
its
affinity
for
and
activity
on
certain
DNA
structures
,
in
a
manner
that
may
enhance
its
relative
specificity
for
physiological
substrates
.
Our
results
also
show
that
acetylation
and
deacetylation
of
endogenous
WRN
is
a
dynamic
process
,
with
sirtuins
and
other
histone
deacetylases
contributing
to
WRN
deacetylation
.
These
findings
advance
our
understanding
of
the
dynamics
of
WRN
acetylation
under
unperturbed
conditions
and
following
DNA
damage
induction
,
linking
this
modification
not
only
to
DNA
damage
persistence
but
also
potentially
to
replication
stalling
caused
by
specific
DNA
lesions
.
Our
results
are
consistent
with
proposed
metabolic
roles
for
WRN
and
genomic
instability
phenotypes
associated
with
WRN
deficiency
.
Diseases
Validation
Diseases presenting
"deficient cells"
symptom
child syndrome
esophageal adenocarcinoma
gm1 gangliosidosis
junctional epidermolysis bullosa
kindler syndrome
lymphangioleiomyomatosis
neonatal adrenoleukodystrophy
werner syndrome
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