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Albinism in phylogenetically and geographically distinct populations of Astyanax cavefish arises through the same loss-of-function Oca2 allele.
[oculocutaneous albinism]
The
Mexican
tetra
,
Astyanax
mexicanus
,
comprises
29
populations
of
cave-adapted
fish
distributed
across
a
vast
karst
region
in
northeastern
Mexico
.
These
populations
have
a
complex
evolutionary
history
,
having
descended
from
'
old
'
and
'
young
'
ancestral
surface-dwelling
stocks
that
invaded
the
region
∼
6
.
7
and
∼
2
.
8
MYa
,
respectively
.
This
study
investigates
a
set
of
captive
,
pigmented
Astyanax
cavefish
collected
from
the
Micos
cave
locality
in
1970
,
in
which
albinism
appeared
over
the
past
two
decades
.
We
combined
novel
coloration
analyses
,
coding
sequence
comparisons
and
mRNA
expression
level
studies
to
investigate
the
origin
of
albinism
in
captive-bred
Micos
cavefish
.
We
discovered
that
albino
Micos
cavefish
harbor
two
copies
of
a
loss
-of-function
ocular
and
cutaneous
albinism
type
II
(
Oca
2
)
allele
previously
identified
in
the
geographically
distant
Pachón
cave
population
.
This
result
suggests
that
phylogenetically
young
Micos
cavefish
and
phylogenetically
old
Pachón
cave
fish
inherited
this
Oca
2
allele
from
the
ancestral
surface-dwelling
taxon
.
This
likely
resulted
from
the
presence
of
the
loss
-of-function
Oca
2
haplotype
in
the
'
young
'
ancestral
surface-dwelling
stock
that
colonized
the
Micos
cave
and
also
introgressed
into
the
ancient
Pachón
cave
population
.
The
appearance
of
albinism
in
captive
Micos
cavefish
,
caused
by
the
same
loss
-of-function
allele
present
in
Pachón
cavefish
,
implies
that
geographically
and
phylogenetically
distinct
cave
populations
can
evolve
the
same
troglomorphic
phenotype
from
standing
genetic
variation
present
in
the
ancestral
taxon
.