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Kindlin-1 controls Wnt and TGF-β availability to regulate cutaneous stem cell proliferation.
[kindler syndrome]
Kindlin-
1
is
an
integrin
tail
binding
protein
that
controls
integrin
activation
.
Mutations
in
the
FERMT-
1
gene
,
which
encodes
for
Kindlin-
1
,
lead
to
Kindler
syndrome
in
man
,
which
is
characterized
by
skin
blistering
,
premature
skin
aging
and
skin
cancer
of
unknown
etiology
.
Here
we
show
that
loss
of
Kindlin-
1
in
mouse
keratinocytes
recapitulates
Kindler
syndrome
and
also
produces
enlarged
and
hyperactive
stem
cell
compartments
,
which
lead
to
hyperthickened
epidermis
,
ectopic
hair
follicle
development
and
increased
skin
tumor
susceptibility
.
Mechanistically
,
Kindlin-
1
controls
keratinocyte
adhesion
through
β
1
-
class
integrins
and
proliferation
and
differentiation
of
cutaneous
epithelial
stem
cells
by
promoting
α
(
v
)
β
(
6
)
integrin-mediated
transforming
growth
factor
-β
(
TGF-β
)
activation
and
inhibiting
Wnt-
β-catenin
signaling
through
integrin-independent
regulation
of
Wnt
ligand
expression
.
Our
findings
assign
Kindlin-
1
the
previously
unknown
and
essential
task
of
controlling
cutaneous
epithelial
stem
cell
homeostasis
by
balancing
TGF-
β-mediated
growth
-inhibitory
signals
and
Wnt-β-catenin-mediated
growth
-promoting
signals
.