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Structural conservation, variability, and immunogenicity of the T6 backbone pilin of serotype M6 Streptococcus pyogenes.
[acute rheumatic fever]
Group
A
streptococcus
(
GAS
;
Streptococcus
pyogenes
)
is
a
Gram
-
positive
human
pathogen
that
causes
a
broad
range
of
diseases
ranging
from
acute
pharyngitis
to
the
poststreptococcal
sequelae
of
acute
rheumatic
fever
.
GAS
pili
are
highly
diverse
,
long
protein
polymers
that
extend
from
the
cell
surface
.
They
have
multiple
roles
in
infection
and
are
promising
candidates
for
vaccine
development
.
This
study
describes
the
structure
of
the
T
6
backbone
pilin
(
BP
;
Lancefield
T
-
antigen
)
from
the
important
M
6
serotype
.
The
structure
reveals
a
modular
arrangement
of
three
tandem
immunoglobulin-like
domains
,
two
with
internal
isopeptide
bonds
.
The
T
6
pilin
lysine
,
essential
for
polymerization
,
is
located
in
a
novel
VAKS
motif
that
is
structurally
homologous
to
the
canonical
YPKN
pilin
lysine
in
other
three
-
and
four
-domain
Gram
-
positive
pilins
.
The
T
6
structure
also
highlights
a
conserved
pilin
core
whose
surface
is
decorated
with
highly
variable
loops
and
extensions
.
Comparison
to
other
Gram
-
positive
BPs
shows
that
many
of
the
largest
variable
extensions
are
found
in
conserved
locations
.
Studies
with
sera
from
patients
diagnosed
with
GAS
-associated
acute
rheumatic
fever
showed
that
each
of
the
three
T
6
domains
,
and
the
largest
of
the
variable
extensions
(
V
8
)
,
are
targeted
by
IgG
during
infection
in
vivo
.
Although
the
GAS
BP
show
large
variations
in
size
and
sequence
,
the
modular
nature
of
the
pilus
proteins
revealed
by
the
T
6
structure
may
aid
the
future
design
of
a
pilus-based
vaccine
.
Diseases
Validation
Diseases presenting
"are promising candidates for vaccine development"
symptom
acute rheumatic fever
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